Clark I don't know what any of this means.
There is a lot of material on the GrapheneOS web site about what things mean and how to use them. With respect to the Network permission, see: https://grapheneos.org/features#network-permission-toggle (which explains why the permission is granted to lots of applications by default).
Clark I have removed almost all permissions from all apps but maybe this will make my device crash?
This is likely to create weird problems that will show up over weeks or months, each one of which will be difficult to diagnose. If you just installed GrapheneOS today, and immediately removed a bunch of permissions from a bunch of system apps, honestly it might be best to factory-reset the device today and start over with a clean slate -- it might save you lots of time later.
By running GrapheneOS, and especially by placing data you care on a device that is running GrapheneOS, you are inherently placing trust in thousands of decisions that the GrapheneOS developers have made over multiple years. Removing permissions from apps that were chosen by the GrapheneOS developers is unlikely to be productive, and is likely to be counterproductive... and meanwhile will not address the acres of non-app code that the GrapheneOS developers also chose (kernel code, system frameworks, etc.).
I'm not suggesting that you must make a leap of faith and 100% trust the GrapheneOS developers without any recourse. The source code is available, so it is possible to see what is happening. If something is wrong, it's possible to build your own copy of GrapheneOS and fix it, and/or submit a fix to the project. This is not a "take it or leave it" system. But it is also not a system that is improved by disabling parts of it without first acquiring deep understanding.
Please note that I do not speak for the GrapheneOS project.